What are the benefits of Spring Framework in Terms of Testing Application?

We have heard a lot about Benefits of Spring like it offers loose coupling, dependency injection, inversion of control etc but from testing point of view, I have couple of question.

• What are the advantages of Spring Framework in terms of testing an application?
• Why Application developed using Spring as considered my testable as opposed to other Web Application?

Please provide some useful examples as then it would be much more easier to understand explanation. I am new to Spring as such and want to understand precise benefits offered by Spring Framework from Application Developer point of view ?